Beetles are considered as primitive pollinators and this has a dual sense. First, among the main pollinator guilds, beetles have been the earliest on the earth to visit systematically flowers and carry over pollen. Second, and as a consequence of the fact that their primeval flower-related characters have little changed since, their primitiveness is recognizable from their body anatomy and their flower-visiting behavior. In the Mediterranean region they are present particularly in the dry months, their massive presence on the flowers denoting the onset of summer drought. The order encompasses generally polyphagous species, therefore species that are not exclusively dependent on flowers.